Tom Shanklin will start in the centre for Wales against England in Cardiff on Saturday after Sonny Parker failed to recover from a trapped neck nerve. Shanklin was first-choice wing in the autumn Tests, but prefers centre and will face England debutant Mathew Tait. Dragons three-quarter Hal Luscombe takes Shanklin's wing berth. Up front, Mefin Davies plays at hooker, open-side Martyn Williams is fit to start and lock Rob Sidoli replaces veteran Gareth Llewellyn. Davies wins the vote ahead of Robin McBryde, who sits on the bench having recently returned to action for the Scarlets following a neck injury. Sidoli is back for the first time since last season's Six Nations, the Blues second row back to form after a niggling groin problem. Williams, Sidoli's team-mate at Cardiff, is set to win his 50th cap, replacing the injured Colin Charvis after making a quicker-than-expected recovery from a neck disc problem. "Initially when I had the diagnosis it was a case of trying to get fit for the Italy game next week, but fortunately my recovery has been a lot quicker than we all hoped," Williams told BBC Sport Wales. "I haven't started a game since 1 January and would have preferred to have a couple of games in the lead-up to this, but the good thing with the injury is that I have kept up my fitness levels. "I feel quite fresh and I just can't wait to play again." Williams says he expects a torrid time from an England loose-forward trio of Lewis Moody, Joe Worsley and Andy Hazell. "They are three top-class players, especially Worsley and Moody. They have been there and done it," Williams added. "They were back-ups behind Richard Hill, Neil Back and Lawrence Dallaglio, but now they have come forward and proved what good players they are - there are no weaknesses there." Luscombe gets the nod on the wing ahead of Dragons team-mate Kevin Morgan, who wins a place on the bench alongside uncapped Blues prop John Yapp. G Thomas (Toulouse (capt); H Luscombe (Dragons), T Shanklin (Blues), G Henson (Ospreys), S Williams (Ospreys); S Jones (Clermont Auvergne), D Peel (Scarlets); G Jenkins (Blues), M Davies (Gloucester), A Jones (Ospreys), B Cockbain (Ospreys), R Sidoli (Blues), D Jones (Scarlets), M Williams (Blues), M Owen (Dragons). R McBryde (Scarlets), J Yapp (Blues), J Thomas (Ospreys), R Jones (Ospreys), G Cooper (Dragons), C Sweeney (Dragons), K Morgan (Dragons).